Toprak Razgatlioglu set the pace early in the 25-minute session on the race tyre, with the Yamaha rider going half a second clear of the field with a 1m39.208s.
th after the first runs had been completed.
Lowes wouldn’t be able to recover much on his qualifying tyre run late in the session and could only manage 14th.
Honda’s Leon Haslam made the first big move on the Q tyre in the final five minutes, lifting his Fireblade up to third briefly.
The reigning British Superbike champion edged ahead of Sykes with a 1m38.736s, though this wasn’t safe as Rea set fastest sector times on his first Q tyre lap.
However, he lost half a second through the third split and called a halt to his charge, leaving Redding to claim his first pole in the series.
Sykes held onto a front row spot in third, 0.040s off Redding, with van der Mark heading row two on the Yamaha.
The Turkish rider will start fourth ahead of Friday pacesetter Ten Loris Baz on the Ten Kate Yamaha, while Chaz Davies (Ducati), Alvaro Bautista (Honda), Haslam and Michael Ruben Rinaldi (Go Eleven Ducati) complete the top 10.
He’ll start 20th in Race 1 this afternoon behind the returning Marco Melandri on the Barni Ducati.